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The reproductive ontogeny of ‘Super-Dwarf’ wheat grown on the space station Mir is chronicled from the vegetative phase through flower' development. Changes in the apical meristem associated with transition from the vegetative plhase to floral initiation and development of the reproductive spike were all typical of ‘Super Dwarf’ wheat up to the point of anthesis. Filament elongation, which characteristically occurs just prior to anthesis (during floral development stage 4) and moves the anthers through the stigmatic branches thus facilitating pollination, did not occur in the flowers of spikes grown on Mir. While pollen did form in the anthers, no evidence of pollination or fertilization was observed. Analysis of pollen idlentified abnormalities; the presence of only one nucleus in the pollen as opposed to the normal trinucleate condition is likely an important factor in the sterility observed in wheat grown on Mir.

A fundamental question for Astrobiology is the question of the ability of life to expand beyond its planet of origin. Introducing life on Mars is the likely near-term step in addressing this question. Making Mars more suitable for life (terraforming) involves altering the martian environment so that microorganisms and plants from Earth could survive there. We define two principal goals: 1) determine the minimal change in pressure, gas composition, and temperature on Mars that would allow for growth of plants from arctic and alpine biomes. 2) Determine the characteristics of plant growth at 0.38 g. This paper reviews martian environmental factors in the context of plant survival, and discusses the use of Space Station as a hypogravity testbed.