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The Mars Exploration Rovers (MER), were launched in June and July of 2003, respectively, and successfully landed on Mars in early and late January of 2004, respectively. The flight system architecture implemented many successful features of the Mars Pathfinder (MPF) system: A cruise stage that transported an entry vehicle that housed the Lander, which in turn, used airbags to cushion the Rover during the landing event. The initial thermal design approach focused on adopting the MPF design wherever possible, and then concentrating on the totally new Rover thermal design. Despite a fundamentally sound approach, there were several salient lessons learned. Some were due to differences from MPF, while others were caused by other means. These lessons sent a clear message: thermal design continues to be a system engineering activity. In each major flight system assembly, there were excellent examples of this recurring theme.