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Proton exchange membrane fuel cell (PEMFC) has attracted extensive attention in recent years because of its high efficiency and zero emission. Although fuel cell technology has made great progress, durability is still the bottleneck of its large-scale commercialization. In order to systematically study the degradation of fuel cell system for transportation applications, we selected the most severe operating condition - on-off cyclic condition as the research object, and focused on the influence of cell temperature and air relative humidity on the degradation rate of fuel cell by means of polarization curve, reference voltage and CV. The results show that with the same temperature, the fuel cell performance degradation rate first decreases and then increases with the decrease of humidity, which may be due to the increase of platinum ion transfer rate at high relative humidity and the dry membrane at low relative humidity.