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The current trend towards an increasing electrification of road vehicles brings to life a whole series of unprecedent design issues. Among these, the ageing process that affects the lifetime of lithium-ion based energy storage systems is of particular importance since it turns out to be extremely sensitive to the variation of battery operating conditions normally occurring especially in hybrid electric vehicles (HEVs). This paper aims at analyzing the impact of operating conditions on the predicted lifetime of a parallel-through-the-road plug-in HEV battery both from thermal and ageing perspectives. The retained HEV powertrain architecture is presented first and modeled, and the related energy management system is implemented. Dedicated numerical models are also discussed for the high-voltage battery pack that allow predicting its thermal behavior and cyclic ageing.